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Tropidoleptus carinatus (Conrad, 1839)

Tropidoleptus carinatus
Tropidoleptus carinatus

Scientific Classification

Kingdom: Animalia
Phylum: Brachiopoda
Class: Rynchonellata
Order: Orthida
Family: Tropidoleptidae
Genus: Tropidoleptus
Species: Tropidoleptus carinatus (Conrad, 1839)

Information

Geological Range

Middle Devonian

Paleogeographic Distribution

Illinois

Indiana

Kentucky

Michigan

Missouri

New York

Ohio

Pennsylvania

West Virginia

Stratigraphic Occurrences

Moscow Formation
Garrattsville member
Windom Member
Kashong member
Deep Run member
Tichenor Member
Ludlowville Formation
Mahantango Formation
Skaneateles Formation
Chenango member
Pompey Member
Delphi Station Member
Mottville Member
Plum Brook Formation
Silica Formation
North Vernon Formation
Swanville Member
Boyle Formation
Lingle Formation
Oatka Creek Formation
Mount Marion Formation

References

Hall (1867): p. 407, pl. 62.

Linsley (1994): p. 171, figs. 10-24.

Wilson (2014): pp. 96, 97.

Remarks

Originally described as Strophomena carinatus by Conrad in 1839. Recombined as Tropidoleptus carinatus by Hall in 1857.

From Wilson (2014, p. 96): “Pedicle valve convex with broad plications and fine growth lines; brachial valve concave. To 20-25mm.”

From Carlton Brett and Gordon Baird: “Tropidoleptus carinatus is an unusual brachiopod. A strophomenid-mimic, the genus has recently been suggested to belong its own order. It is a long ranging species that spans much of the Devonian Period but was clearly not a member of the Eastern Americas Realm and arrived as a part of a wave of immigration of Old World Realm taxa associated with the incoming of the Hamilton fauna and the Kacak global bioevent.; T. carinatus is widespread in nearshore siliciclastic facies of the Appalachian basin, often associated with Mucrospirifer and varied bivalves in siltstone-sandstone facies. It is rare or absent from Diverse brachiopod (e.g., Pseudoatrypa-Megastrophia associations) but occurs abundantly in laterally equivalent mudstone facies with a distinctive suite of other taxa including Devonochonetes coronatus (also an apparent immigrant taxon) and Nucleospira, bryozoans and diverse echinoderms and the domal tabulate Pleurodictyum but generally lacking rugose corals, in beds swirled by Zoophycos, suggestive of high turbidity. This Tropidoleptus-Nucleospira biofacies is best exemplified by the Kashong Shale (Moscow Formation) but also occurs in shallow parts of cycles throughout the Hamilton Group. This brachiopod is commonly encrusted, particularly by Hederella and microconchids. A peculiarity of this species is its abrupt abundant appearance together with D. coronatus in the early Givetian lower Skaneateles Mottville (NY), Rockport Quarry (Ontario subsurface), lower Silica (northern Ohio), and Swanville (IN) formations. This faunal epibole is a useful stratigraphic marker. Curiously, in these units these normally siliciclastic associated brachiopods occur in shallow water carbonate grainstones.; In addition, there are some beds of medium to dark gray shale with low diversity assemblages with other diminutive brachiopods including Cruspina nana, Truncalosia, small Rhipidothyris or even tiny Allanella (see discussion under those species) in the Ludlowville and Moscow formations typical of dysoxic facies, in which bedding planes are covered by diminutive specimens of Tropidoleptus. These specimens which rarely exceed 10 mm resemble small (juvenile) individuals in normal T. carinatus populations. However, they do not form the lower end of a gradient, rather they are a discrete biofacies. It is unclear whether these represent all failed spatfalls with juvenile mortality or rather a small perhaps paedomorphic form of Tropidoleptus.”

Online Resources

Paleobiology Database.

Media

Images

Photograph of fossil of Tropidoleptus carinatus (PRI 109484).
Tropidoleptus carinatus from the Ludlowville Fm. (Wanakah Mbr.) of Erie County, New York, With crinoid holdfast on shell (PRI 109484).
Photograph of fossil of Tropidoleptus carinatus (PRI 109485).
Tropidoleptus carinatus from the Ludlowville Fm. (Wanakah Mbr.) of Erie County, New York (PRI 109485).
Photograph of fossil of Tropidoleptus carinatus (MPM P11080).
Tropidoleptus carinatus from the Devonian of Bethany, New York (stratigraphic position uncertain) (MPM P11080).
Photograph of fossil of Tropidoleptus carinatus (MPM P11080).
Tropidoleptus carinatus from the Devonian of Bethany, New York (stratigraphic position uncertain) (MPM P11080).
Illustrations from Hall of fossils of Tropidoleptus carinatus.
Illustrations of Tropidoleptus carinatus from Hall, reproduced in Linsley (1994, p.171, figs 10-24) and Wilson (2014, pp. 96, 97).

Related

Rhipidomella leucosia
Rhipidomella penelope
Rhipidomella vanuxemi